Diversity of fish
Fish are very diverse and are categorized in many ways. This article is an overview of some of the more common types of fish. Although most fish species have probably been discovered and described, about 250 new ones are still discovered every year. According to FishBase, 31,500 species of fish had been described by January 2010. That is more than the combined total of all other vertebrates: mammals, amphibians, reptiles and birds.
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[edit] By species
Fish systematics is the formal description and organisation of fish taxa into systems. It is complex and still evolving. Controversies over "arcane, but important, details of classification are still quietly raging."
The term "fish" describes any non-tetrapod chordate, (i.e., an animal with a backbone), that has gills throughout life and has limbs, if any, in the shape of fins. Unlike groupings such as birds or mammals, fish are not a single clade but a paraphyletic collection of taxa, including jawless, cartilaginous and skeletal types.
[edit] Jawless fish
Jawless fish are the most primitive fish. There is current debate over whether these are really fish at all. They have no jaw, no scales, no paired fins, and no bony skeleton. Their skin is smooth and soft to the touch, and they are very flexible. Instead of a jaw, they possess an oral sucker. They use this to fasten on to other fish, and then use their rasp-like teeth to grind through their host's skin into the viscera. Jawless fish inhabit both fresh and salt water environments. Some are anadromous, moving between both fresh and salt water habitats.
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Extant jawless fish are either lamprey or hagfish. Juvenile lamprey feed by sucking up mud containing micro-organisms and organic debris. The lamprey has well developed eyes, while the hagfish has only primitive eyespots. The hagfish coats itself and carcasses it finds with noxious slime to deter predators, and periodically ties itself into a knot to scrape the slime off. It is the only invertebrate fish and the only animal which has a skull but no vertebral column. It has four hearts, two brains, and a paddle-like tail.[[1]]
[edit] Cartilaginous fish
Cartilaginous fish have a cartilaginous skeleton. However, their ancestors were bony animals, and were the first fish to develop paired fins. Cartilaginous fish don't have swim bladders. Their skin is covered in denticles, that are as rough as sandpaper. Because cartilaginous fish do not have bone marrow, the spleen and special tissue around the gonads produces red blood cells. Some cartilaginous fishes possess an organ called Leydig's Organ which also produces red blood cells.
| There are over 980 species of cartilaginous fish. They include sharks, rays and chimaera. | |||
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[edit] Bony fish
Bony fish include the lobe finned fish and the ray finned fish. The lobe finned fish is the class of fleshy finned fishes, consisting of lungfish, and coelacanths. They are bony fish with fleshy, lobed paired fins, which are joined to the body by a single bone. These fins evolved into the legs of the first tetrapod land vertebrates, amphibians. They are called ray finned because they possess lepidotrichia or "fin rays", their fins being webs of skin supported by bony or horny spines ("rays").
There are three types of ray finned fishes: the chondrosteans, holosteans, and teleosts. The chondrosteans and holosteans are primitive fishes sharing a mixture of characteristics of teleosts and sharks. In comparison with the other chondrosteans, the holosteans are closer to the teleosts and further from sharks.
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[edit] Teleosts
Teleosts are the most advanced or "modern" fishes. They are overwhelmingly the dominant class of fishes (or for that matter, vertebrates) with nearly 30,000 species, covering about 96 percent of all extant fish species. They are ubiquitous throughout fresh water and marine environments from the deep sea to the highest mountain streams. Included are nearly all the important commercial and recreational fishes.[[2]]
Teleosts have a movable maxilla and premaxilla and corresponding modifications in the jaw musculature. These modifications make it possible for teleosts to protrude their jaws outwards from the mouth. The caudal fin is homocercal, meaning the upper and lower lobes are about equal in size. The spine ends at the caudal peduncle, distinguishing this group from those in which the spine extends into the upper lobe of the caudal fin.
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[edit] By size
The smallest species is Paedocypris progenetica, a type of minnow. It is also the smallest vertebrate. The smallest mature female had a standard length of 7.9 millimeters (0.3 in). The largest individual is 10.3 millimeters (0.4 in). They live in the dark–colored peat swamps of the Indonesian island of Sumatra.[[3]]
Other contenders for smallest fish are the male anglerfish, Photocorynus spiniceps,[[4]] and the stout infantfish, a type of goby. [[5]]According to the Guinness Book of World Records, the sinarapan, another type of goby, is the world's smallest commercially harvested fish. Found in the Philippines, they have an average length of 12.5 millimeters (0.5 in), and are threatened by overfishing.
The largest is the whale shark. It is a slow moving filter feeding shark with a maximum published length of 20 meters (66 ft) and a maximum weight of 34 tonnes. Whale sharks can live up to 70 years.[[6]]
The heaviest bony fish is the ocean sunfish. It can weigh up to 2,300 kg (5,100 lb). It is found in all warm and temperate oceans. The longest bony fish is the king of herrings. Its total length can reach 11 metres (36 ft), and it can weigh up to 272 kilograms (600 lb). It is a rarely seen oarfish found in all the world's oceans, at depths of between 20 metres (66 ft) and 1,000 metres (3,300 ft).[[7]]
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[edit] By life span
Some of the shortest-lived species are gobies, which are small coral reef–dwelling fish. Some of the longest-lived are the rockfish.
The shortest lived is the seven-figure pygmy goby, which lives for at most 59 days. This is the shortest lifespan for any vertebrate.[[8]] Short lived fish have particular value in genetic studies on aging. In particular, the ram cichlid is used in laboratory studies because of its ease of breeding and predictable aging pattern.[[9]] [[10]]
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The longest–lived fish is the 205 years reported for the rougheye rockfish, Sebastes aleutianus, found offshore in the North Pacific at 25–900 metres (14–490 fathoms). This fish exhibits negligible senescence.[[11]] [[12]]
The longest living commercial fish may be the orange roughy, with a maximum reported age of 149 years.[[13]] One of the longest living sport fish is the Atlantic tarpon, with a maximum reported age of 55 years."[[14]] [[15]] [[16]]
There are stories about Japanese Koi goldfish passed from generation to generation for 300 years. Scientists are sceptical. Counting growth lines on the scales of fish confined to ponds or bowls is unreliable, since they lay down extra lines.[[17]] [[18]] The maximum reliably reported age for a goldfish is 41 years.[[19]]
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Some of the longest living fish are living fossils, such as the green sturgeon. These species are among the longest living species found in freshwater, with a maximum reported age of 60 years. They are also among the largest fish species found in freshwater, with a maximum reported length of 2.5 meters (8 ft) and a maximum reported weight of {{kg to lb}159}}. [[20]] [[21]]Another living fossil is the Australian lungfish. One individual has lived in an aquarium for 75 years, and is the oldest fish in captivity. According to fossil records, the Australian lungfish has hardly changed for 380 million years.[[22]] [[23]] [[24]]
[edit] By habitat
There is 10,000 times more saltwater in the oceans than there is freshwater in the lakes and rivers. However, only 58 percent of extant fish species are saltwater. A disproportionate 41 percent are freshwater fish (the remaining one percent are anadromous). This diversity in freshwater species is, perhaps, not surprising, since the thousands of separate lake habitats promote speciation.
| Habitat | Area | Volume | Depth | Species | Fish biomass | |
|---|---|---|---|---|---|---|
| million km2 | million cu km | (mean) | count | percent | million tonnes | |
| Saltwater | 361 | 1370.8 | 3.8km | 18,000 | 58 | 800-2,000 |
| Freshwater | 1.5 | 0.13 | 87m | 13,000 | 41 | |
Fish can also be demersal or pelagic. Demersal fish live on or near the bottom of oceans and lakes, while pelagic fish inhabit the water column away from the bottom. Habitats can also be vertically stratified. Epipelagic fish occupy sunlit waters down to 200 metres (110 fathoms), mesopelagic fish occupying deeper twilight waters down to 1,000 meters (3,281 ft), and bathypelagic fish inhabiting the cold and pitch black depths below.
Most oceanic species (78 percent, or 44 percent of all fish species), live near the shoreline. These coastal fish live on or above the relatively shallow continental shelf. Only 13 percent of all fish species live in the open ocean, off the shelf. Of these, 1 percent are epipelagic, 5 percent are pelagic, and 7 percent are deep water.
Fish are found in nearly every aquatic habitat. Most fish, whether by species count or abundance, live in warmer environments with relatively stable temperatures. However, some species can survive temperatures up to 44.6 °C (112.3 °F), while others can cope with salinity greater than 10 percent. The world's deepest living fish, Abyssobrotula galatheae, a species of cusk eel, lives in the Puerto Rico Trench at a depth of 8,372 meters (27,467 ft).[[25]] At the other extreme, the Tibetan stone loach lives at altitudes over 5,200 meters (17,060 ft) in the Himalayas.[[26]]
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Some marine pelagic fish range over vast areas, such as the blue shark that lives in all oceans. At the other extreme are fish confined to single, small living spaces, such as isolated cave fish like Lucifuga in the Bahamas and Cuba, or equally isolated desert pupfish living in small desert spring systems in Mexico and the southwest U.S., or bythitid vent fish like Thermichthys hollisi, living around thermal vents 2,400 metres (1,300 fathoms) down.[[27]]
[edit] By breeding behavior
Grouper are protogynous hermaphrodites, who school in harems of three to fifteen females. When no male is available, the most aggressive and largest females shift sex to male, probably as a result of behavioral triggers.
In very deep waters, it is not easy for a fish to find a mate. There is no light, so some species depend on bioluminescence. Others are hermaphrodites, which doubles their chances of producing both eggs and sperm when an encounter does occur.[[28]] The female anglerfish releases pheromones to attract tiny males. When a male finds her, he bites on to her and never lets go. When a male of the anglerfish species Haplophryne mollis bites into the skin of a female, he release an enzyme that digests the skin of his mouth and her body, fusing the pair to the point where the two circulatory systems join up. The male then atrophies into nothing more than a pair of gonads. This extreme sexual dimorphism ensures that, when the female is ready to spawn, she has a mate immediately available.[[29]]
Some sharks, such as hammerheads are able to breed parthogenetically.
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[edit] By brooding behavior
Fish adopt a variety of strategies for nurturing their brood. Sharks, for example, variously follow three protocols with their brood. Most sharks, including lamniformes [[30]]are ovoviviparous, bearing their young after they nourish themselves after hatching and before birth, by consuming the remnants of the yolk and other available nutrients. Some such as hammerheads [[31]] are viviparous, bearing their young after nourishing hatchlings internally, analgously to mammalian gestation. Finally catsharks [[32]] and others are, oviparous, laying their eggs to hatch in the water.
Some animals, predominantly fish such as cardinalfish [[33]] practice mouthbrooding, caring for their offspring by holding them in the mouth of a parent for extended periods of time. Mouthbrooding has evolved independently in several different families of fish.
Others, such as seahorse males, practice pouch-brooding, analogous to Australia's kangaroos, nourishing their offspring in a pouch in which the female lays them. [[34]]
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[edit] By feeding behavior
There are three basic methods by which food is gathered into the mouths of fish: by suction feeding, by ram feeding, and by manipulation or biting. [[35]]Nearly all fish species use one of these styles, and most use two.
Early fish lineages had inflexible jaws limited to little more than opening and closing. Modern telosts have evolved protusible jaws that can reach out to engulf prey. An extreme example is the protusible jaw of the slingjaw wrasse. Its mouth extends into a tube half as long as its body, and with a strong suction it catches prey. The equipment tucks away under its body when it is not in use.
In practice, feeding modes lie on a spectrum, with suction and ram feeding at the extremes. Many fish capture their prey using both suction pressure combined with a forward motion of the body or jaw.[[36]]
The cookiecutter shark is a small dogfish which derives its name from the way it removes small circular plugs, looking as though cut with a cookie cutter, from the flesh and skin of cetaceans and larger fish, including other sharks. The cookiecutter attaches to its larger prey with its suctorial lips, and then protrudes its teeth to remove a symmetrical scoop of flesh.[[37]]
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Most fish are food opportunists, or generalists. They eat whatever is most easily available. For example, the blue shark feeds on dead whales and nearly everything else that wriggles: other fish, cephalopods, gastropods, ascidians, crustaceans.[[38]] Ocean sunfish prefer jellyfish.
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Other fish have developed extreme specializations. Silver arowana, also called monkey fish, can leap two meters out of the water to capture prey. They usually swim near the surface of the water waiting for potential prey. Their main diet consist of crustaceans, insects, smaller fishes and other animals that float on the water surface, for which its draw-bridge-like mouth is exclusively adapted for feeding. The remains of small birds, bats, and snakes have also been found in their stomachs.[[39]]
Archerfish prey on land–based insects and other small animals by literally shooting them down with water droplets from their specialized mouths. Archerfish are remarkably accurate; adults almost always hit the target on the first shot. They can bring down insects such as grasshoppers, [[40]]spiders and butterflies on a branch of an overhanging tree,[[41]] 3 m above the water's surface.[[42]] This is partially due to good eyesight, but also the ability to compensate for light refraction when aiming.[[43]] Triggerfish also use jets of water, to uncover sand dollars buried in sand or overturn sea urchins.
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Doctor fish (nibble fish) live and breed in the outdoor pools of some Turkish spas, where they feed on the skin of patients with psoriasis. The fish are like cleaner fish in that they only consume the affected and dead areas of the skin, leaving the healthy skin to recover.
[edit] By swimming ability
The slowest-moving fishes are the sea horses. The slowest of these, the dwarf seahorse, attains about five feet per hour.
The fasted sprinter is the Indo-Pacific sailfish. It has been recorded in a burst of over 110 kilometres per hour (68 mph), equivalent to 12 to 15 times its own length per second. The wahoo is perhaps the fastest fish for its size, attaining a speed of 19 lengths per second, reaching 78 kilometres per hour (48 mph).
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The shortfin mako shark is fast enough and agile enough to chase down and kill an adult swordfish, but they don't always win. Sometimes in the struggle with a shark a swordfish can kill it by ramming it in the gills or belly. The shortfin mako's speed has been recorded at 50 kilometres per hour (31 mph), and there are reports that it can achieve bursts of up to 74 kilometres per hour (46 mph). It can jump up to 9 meters (30 ft) in the air. Due to its speed and agility, this high-leaping fish is sought as game worldwide. This shark is highly migratory. Its exothermic constitution partly accounts for its relatively great speed.
The northern bluefin tuna is capable of sustained high speed cruising, and maintains high muscle temperatures so it can can cruise in arctic waters.
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A number of species jump while swimming near the surface, skimming the water. Although not exactly a "swimming" ability, flying fish have unusually large pectoral fins, which enable the fish to take short gliding flights above the surface of the water, in order to escape from predators. Their glides are typically around 50 meters (160 ft), but they can use updrafts at the leading edge of waves to cover distances of at least 400 meters (1,300 ft). In May 2008, a flying fish was filmed off the coast of Japan (see video). The fish spent 45 seconds aloft, and was able to stay aloft by occasionally beating the surface of the water with its caudal (tail) fin. The previous record was 42 seconds.
[edit] By toxicity
Toxic fish produce strong poisons in their bodies. Both poisonous fish and venomous fish, contain toxins, but deliver them differently.
Venomous fish bite, sting, or stab, causing an envenomation. Venomous fish don't necessarily cause poisoning if they are eaten, since the digestive system often destroys the venom. By contrast, the digestive system does not destroy poisonous fish toxins, making them poisonous to eat.[[44]]
The most poisonous fish is the puffer fish. It is the second most poisonous vertebrate after the golden dart frog. It paralyzes the diaphragm muscles of human victims, who can die from suffocation. In Japan, skilled chefs use parts of a closely related species, the blowfish to create a delicacy called "fugu", including just enough toxin for that "special flavour".
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The spotted trunkfish is a reef fish which secretes a colourless ciguatera toxin from glands on its skin when touched. The toxin is only dangerous when ingested, so there's no immediate harm to divers. However, predators as large as nurse sharks can die as a result of eating a trunkfish.
The giant moray is a reef fish at the top of the food chain. Like many other apex reef fish, it is likely to cause ciguatera poisoning if eaten. Outbreaks of ciguatera poisoning in the 11th to 15th centuries from large, carnivorous reef fish, caused by harmful algal blooms, could be a reason why Polynesians migrated to Easter Island, New Zealand, and possibly Hawaii.
A 2006 study found that at least 1200 species of venomous fish. There are more venomous fish than venomous snakes. In fact, there are more venomous fish than the combined total of all other venomous vertebrates. Venomous fish are found in almost all habitats around the world, but mostly in tropical waters. They wound over 50,000 people every year.
They carry their venom in venom glands and use various delivery systems, such as spines or sharp fins, barbs, spikes and fangs. Venomous fish tend to be either very visible, using flamboyant colors to warn enemies, or skilfully camouflaged and maybe buried in the sand. Apart from the defense or hunting value, venom help bottom dwelling fish by killing the bacteria that try to invade their skin. Few of these venoms have been studied. They are yet to be tapped resource for bioprospecting to find drugs with medical uses.
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The most venomous fish is the reef stonefish. It has a remarkable ability to camouflage itself amongst rocks. It is an ambush predator that sits on the bottom waiting for prey to approach. Instead of swimming away if disturbed, it erects 13 venomous spines along its back. For defense, it can shoot venom from each or all of these spines. Each spine is like a hypodermic needle, delivering the venom from two sacs attached to the spine. The stonefish has control over whether to shoot its venom, and does so when provoked or frightened. The venom results in severe pain, paralysis and tissue death, and can be fatal if not treated. Despite its formidable defenses, stonefish have predators. Some bottom feeding rays and sharks with crushing teeth feed on them, as does the Stokes' seasnake.
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Unlike stonefish, lionfish can only release venom when something strikes its spines. Although not native to the U.S. coast, lionfish have appeared around Florida and have spread up the coast to New York. They are attractive aquarium fish, sometimes used to stock ponds, and may have been washed into the sea during a hurricane. Lionfish can aggressively dart at scuba divers and attempt to puncture their facemask with their venomous spines.
The stargazer buries itself and can deliver electric shocks as well as venom. It is a delicacy in some cultures (cooking destroys the venom), and can be found for sale in some fish markets with the electric organ removed. They have been called "the meanest things in creation"
Stingray envenomations can occur to people who wade in shallow water and tread on them. This can be avoided by shuffling through the sand or stamping on the bottom, as the rays detect this and swim away. The stinger usually breaks off in the wound. It is barbed, so it can easily penetrate but not so easily be removed. The stinger causes local trauma from the cut itself, pain and swelling from the venom, and possible later infection from bacteria. Occasionally severed arteries or death can result.
Treatment for venom stings usually includes the application of heat, using water at temperatures of about 45 °C (113 °F), since heat breaks down most complex venom proteins.
[edit] By human use
Fish are sought by humans for their value as commercial food fish, recreational sport fish, decorative aquarium fish and in tourism, attracting snorkelers and SCUBA divers .
Throughout human history, important fisheries have been based on forage fish. Forage fish are small, schooling for protection which are eaten by larger predators. Typical ocean forage fish feed near the bottom of the food chain on plankton, often by filter feeding. They include the family Clupeidae (herrings, sardines, menhaden, hilsa, shad and sprats), as well as anchovies, capelin and halfbeaks.
Important herring fisheries have existed for centuries in the North Atlantic and the North Sea. Likewise, important traditional for anchovy and sardine fisheries have operated in the Pacific, the Mediterranean, and the southeast Atlantic. The world annual catch of forage fish in recent years has been around 25 million tonnes, or one quarter of the world's total catch.
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Higher in the food chain, Gadidae (cod, pollock, haddock, saithe, hake and whiting also support important fisheries. Concentrated initially in the North Sea, Atlantic cod was one of Europe's oldest fisheries, later extending to the Grand Banks, Declining numbers led to international "cod wars" and eventually the virtual abandonment of these fisheries. These days the Alaska pollock supports an important fishery in the Bering Sea and the north Pacific, yielding about 6 million tonnes, while cod amounts to about 9 million tonnes.
- Food fish, Oily fish, whitefish
- Farmed fish
- Fish used for medicinal purposes
Recreational and sport fishing is big business[108] U.S. saltwater fishers spend about $30 billion annually and support 350,000 jobs. Some of the more popular recreational and sport fish include bass, marlin, porgie, shad, mahi-mahi, smelt whiting, swordfish, and walleye.
Fishkeeping is another popular pastime, and there is a large international trade for aquarium fish.
Snorkeling and SCUBA diving attract millions of people to beaches, coral reefs, lakes, and other water bodies to view fish and other marine life.
[edit] By vulnerability
- resilience
[edit] Other types
Fish hold the records for the relative brain weights of vertebrates. Most vertebrate species have similar brain-to-body weight ratios. The deep sea bathopelagic cusk-eel Acanthonus armatus, has the smallest ratio of all known vertebrates. At the other extreme, the elephantnose fish, an African freshwater fish, has the largest ratio of all known vertebrates.
Sarpa salpa, a species of bream are recognizable by the golden stripes running the length of its body and can induce LSD-like hallucinations if it is eaten. These widely distributed coastal fish became a recreational drug during the Roman Empire, and are called "the fish that make dreams" in Arabic. Other hallucinogenic fish are Siganus spinus, called "the fish that inebriates" in Reunion Island, and Mulloidichthys samoensis, called "the chief of ghosts" in Hawaii.[[45]]
[edit] External links
- Articles About Marine Life Oceans for Youth Foundation.
